What is the difference between PCIe Gen 3 and PCIe Gen 4?

What is the difference between PCIe Gen 3 and PCIe Gen 4? nvme gen 4 in gen 3 slot
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nvme in pcie slot 1x SSD slot , 1x SSD Combo (NVMe PCIe Gen3 1x 2280 NVMe Gen 3 slot + 1x 2280 Gen 4 ; 1x 2280 NVMe Gen

nvme pcie 40 in 30 slot I was wondering if I could use Nvme gen4 on a gen 3 board ? I think my board only has one slot, so I may want to upgrade 5+3 Phases Pure Digital VRM Solution with Low RDS MOSFETs 9 Supports 3rd Gen AMD Ryzen™ Processors 10 Dual Channel DDR4, 4 DIMMs 11 Dual